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When did haplogroup R-Z16502 mutate from FGC5494?

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Haplogroup R-Z16502. My family arrived in Accomack Co, VA in 1636. I'm curious when haplogroup mutation occurred.  4,000 years ago; before or after 1066 AD? etc.  Family Tree DNA indicates Mutation path: R-M269>P311>P312>L21>FGC5494>R-Z16502.
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in Genealogy Help by Richard Colbert G2G2 (2.5k points)

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